When a group of people capture Penryn's sister Paige, thinking she's a monster, the situation ends in a massacre. Paige disappears. Humans are terrified. Mom is heartbroken.
Penryn drives through the streets of San Francisco looking for Paige. Why are the streets so empty? Where is everybody? Her search leads her into the heart of the angels' secret plans where she catches a glimpse of their motivations, and learns the horrifying extent to which the angels are willing to go.
Meanwhile, Raffe hunts for his wings. Without them, he can't rejoin the angels, can't take his rightful place as one of their leaders. When faced with recapturing his wings or helping Penryn survive, which will he choose?